Is Gender Identity Unique to Humans?

This summer, in the introductory course I teach on the evolution and biology of human and animal behavior, I showed my students a website that demonstrates how to identify frog “genders.” I explained that this was a misuse of the term “gender”; what the author meant was how to identify frog sexes. Gender, I told the students, goes far beyond mere sex differences in appearance or behavior. It refers to something complex and abstract that may well be unique to Homo sapiens.